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Although, no proof for sulfenic acid formation was provided, the authors speculated that Cys511 of Cys511Cys535 disulfide first reacted with hydroxide ion to expel the Cys535 thiolate, which in turn, reacted with another disulfide to give disulfide scrambling

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Webster TMU, Santos EM (2015) Global transcriptomic profiling demonstrates induction of oxidative stress and of compensatory cellular stress responses in brown trout exposed to glyphosate and Roundup