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Funding This work was supported by the National Key R&D Program of China (2019YFD1001300, 2019YFD1001303), the Critical Talent Workstation Project (TYSGJ202201), the National Natural Science Foundation of China (31960415), the Academician Workstation Project (TYYSZ201707), the Research Program Sponsored by State Key Laboratory of Integrative Sustainable Dryland Agriculture, Shanxi Agricultural University 202105D121008-3-5), the Fundamental Research Project of Shanxi Province (Grant No