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PubMed: (peer-reviewed) Lee S, et al

Analysis of the PPARGC1A promoter sequence up to 2,000 bp before the transcription initiation site (Genomatix) and sequence comparison between the human and murine PPARGC1A promoter sequence (CLUSTAL O 1.2.0 sequence alignment software) (Supplemental Figure 4A) identified 5 conserved PPREs that span regions 1631/1609 bp, 1386/1362 bp, 1012/991 bp, 634/612 bp, and 210/189 bp (Supplemental Figure 4B)

Importantly, nuclear-localized FOXO4-positive Leydig cells in elderly testes showed reduced expression of 3-HSD, a rate-limiting enzyme in testosterone synthesis, implying a connection between nuclear FOXO4 and age-related decline in testosterone production [1]