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In starch synthesis, AGP catalyzes the conversion of glucose 1-phosphase and ATP to produce the precursor for starch biosynthesis, ADP-glucose (Sun et al., 2021), while GBSS and SSS catalyze the elongation of a-1,4-glucosidic bonds on amylose and amylopectin molecules, respectively, by the transfer of glucose from ADPG (Wu et al., 2020)