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Excessive Sun Exposure Prolonged exposure to UV rays triggers melanin production, leading to lip darkening

This upregulation leads to an increased expression of lipogenic genes, including sterol regulatory element-binding proteins (SREBP1c) and diacylglycerol acyltransferase-2 (DGAT2), which suggests that miR-370 provides a further point of regulation of this pathway 86

Garber SB, Kingsbury JS, Gray BL, Hoveyda AH (2000) Efficient and recyclable monomeric and dendritic Ru-Based metathesis catalysts

Dark spots or hyperpigmentation, including those left behind after acne heals (called post-inflammatory hyperpigmentation), are caused by excess melanin